In Heian Castle, humans, raccoons, and tengu are in a three-way tripod, turning the huge wheel of the city together. Shimogamo Yasaburo (voiced by Sakurai Takahiro) is the third son of the famous Shimogamo family in the raccoon world. Although he is a raccoon, he can transform into a human at will. His mentor is Tengu Akadama. The old Akadama's magic power is not as good as before, and he is in love with the beautiful human woman Benten (voiced by Noto Mamiko). Benten is beautiful and unruly by nature. She is a member of the secret organization "Friday Club" in Kyoto. The owner of the Shimogamo family, Soichiro, was deeply admired by the raccoon family, but he passed away several years ago, and the story remains a mystery. He left behind four unsuccessful sons: the eldest son Shimogamo Yaichiro (voiced by Suwabe Junichi) is old-fashioned and indecisive; the second son Yajiro is introverted and autistic, not asking about worldly affairs; Yasaburo only cares about fun in everything; the youngest brother Yashiro is as timid as a mouse. This is the Shimogamo family living in the forest of the knot. Their mother is convinced that one day, her four children will inherit their late father's legacy and become great raccoon cats...