The battle between the Autobots and the Decepticons has lasted for centuries and has never ended. Their descendants, the Colossus and the Protobeast, have pushed a new round of battles into the prehistoric era. On the way to find new energy sources, the Tyrannosaurus Rex (voiced by David Kaye) commanded the Protobeast to jump through time and space to the prehistoric Earth, and the Colossus followed under the leadership of Optimus Prime (voiced by Garry Chalk). The tug-of-war between the two sides continued, and soon even the alien Walkers who had long been entrenched on the planet were involved. The information about the future carried by the time traveler frightened the aliens, making the battle more intense. After that, the Tyrannosaurus Rex discovered the Ark, a battleship left on Earth by the Autobots. In order to change the history of the Decepticons' defeat, they tried to destroy the sleeping Optimus Prime. The battle that concerns the fate of the entire universe kicked off in advance in ancient times...